1. College of Life Sciences & Medicine, School of Biological Sciences, Zoology Building; University of Aberdeen; Tillydrone Avenue Aberdeen AB24 2TZ U.K.
2. Faculty of Biosciences, Economy and Fisheries, Department of Arctic and Marine Biology; University of Tromsø; 9037 Tromsø Norway
3. Marine Biological Association of the United Kingdom, The Laboratory, Plymouth, PL1 2PB, U.K. and School of Environment and Life Sciences, University of Salford, Greater Manchester M5 4WT; U.K.