1. Disease Resistant Crops Research Unit National Institute of Agrobiological Sciences 2‐1‐2 Kannondai Tsukuba Ibaraki 305‐8602 Japan
2. Graduate School of Life and Environmental Sciences University of Tsukuba Tsukuba Ibaraki Japan
3. International Center for Tropical Agriculture (CIAT) Cali Colombia
4. Agrogenomics Research Center National Institute of Agrobiological Sciences Tsukuba Ibaraki Japan