1. State Key Laboratory of Marine Environmental Science and College of Ocean and Earth Sciences Xiamen University Xiamen, Fujian China
2. Department of Marine Sciences University of Connecticut Groton Connecticut USA
3. State Key Laboratory of Marine Resource Utilization in South China Sea Hainan University Haikou, Hainan China
4. Laboratory of Marine Biology and Biotechnology Qingdao National Laboratory of Marine Science and Technology Qingdao, Shandong China