1. Innovation Platform of Molecular Biology, College of Landscape and Art Henan Agricultural University Zhengzhou 450046 China
2. Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, School of Life Sciences Fudan University Shanghai 200438 China
3. U.S. Forest Service Rocky Mountain Research Station 2500S. Pine Knoll Dr. Flagstaff Arizona USA
4. School of Life Sciences Henan University Kaifeng 475001 Henan China
5. Key Laboratory of Stress Physiology and Ecology in Cold and Arid Regions, Department of Ecology and Agriculture Research, Northwest Institute of Eco‐Environment and Resources Chinese Academy of Sciences Lanzhou 730000 China
6. South China Botanical Garden Chinese Academy of Sciences Guangzhou 510650 China