1. Department of Fisheries; Faculty of Fisheries and Environmental Sciences; Gorgan University of Agricultural Sciences and Natural Resources; Gorgan Iran
2. Razi Vaccine and Serum Research Institute; Agricultural Research, Education and Extension Organization (AREEO); Karaj Iran
3. Faculty of Bioscience, Fisheries and Economics; Norwegian College of Fishery Science; UiT The Arctic University of Norway; Breivika Norway