1. Department of Biology; Centre for Biodiversity Dynamics; Norwegian University of Science and Technology; Trondheim Norway
2. School of Biology and Center of Excellence in Phylogeny of Living Organisms; University of Tehran; Tehran Iran
3. Department of Biology; Norwegian University of Science and Technology; Trondheim Norway
4. School of Biological Sciences; King Henry Building; University of Portsmouth; Portsmouth UK
5. Institute of Arctic Biology; University of Alaska; Fairbanks AK USA