1. Department of Botany, Ecology and Plant Physiology University of La Laguna (ULL) Tenerife Spain
2. Department of Plant Biology and Ecology University of the Basque Country (UPV/EHU) Leioa Spain
3. Organismal and Evolutionary Biology (OEB), Viikki Plant Science Centre (ViPS) Faculty of Biological and Environmental Sciences, University of Helsinki Helsinki Finland