1. Department of Biology University of Hawaiʻi at Hilo Hilo HI 96720 USA
2. Department of Botany, MRC166 Smithsonian Institution P.O. Box 37012 Washington DC 20013‐7012 USA
3. USDA‐ARS Daniel K. Inouye U.S. Pacific Basin Agricultural Research Center 64 Nowelo St. Hilo HI 96720 USA
4. National Tropical Botanical Garden 3530 Papalina Road, Kalāheo Kauaʻi HI 96741 USA
5. Department of Ecology & Evolutionary Biology Biodiversity Institute University of Kansas Lawrence KS 66045‐7534 USA
6. Laboratorio de Botánica, Departamento de Biología Vegetal, Facultad de Agronomía Universidad de la República Av. Garzón 780 Montevideo CP 12900 Uruguay
7. School of Life Sciences University of Hawaiʻi at Mānoa Honolulu HI 96822 USA
8. Délégation à la Recherche Government of French Polynesia Papeete Tahiti French Polynesia
9. Santa Barbara Botanic Garden, Herbarium Santa Barbara Botanic Garden 1212 Mission Canyon Road Santa Barbara CA 93105 USA