1. Key Laboratory of Biodiversity and Biogeography; Kunming Institute of Botany; Chinese Academy of Sciences; Kunming 650204 China
2. Laboratory of Subtropical Biodiversity; Jiangxi Agricultural University; Nanchang Jiangxi 330045 China
3. School of Life Science; Yunnan Normal University; Kunming 650092 Yunnan China
4. Department of Organismic Biology; University of Salzburg; Hellbrunnerstrasse 34 A-5020 Salzburg Austria