1. Key Laboratory of Tropical Marine Bio-resources and Ecology (LMB); Key Laboratory of Applied Marine Biology of Guangdong Province and Chinese Academy of Sciences (LAMB); South China Sea Institute of Oceanology; Chinese Academy of Sciences; Guangzhou China
2. Tropical Marine Biological Research Station in Hainan; South China Sea Institute of Oceanology; Chinese Academy of Sciences; Sanya China
3. School of Pharmaceutical Sciences; South Medical University; Guangzhou China
4. Department of Aquaculture; College of Animal Science; South China Agricultural University; Guangzhou China