1. Quantitative and Applied Ecology Group, School of BioSciences; University of Melbourne; Parkville 3010 Melbourne Victoria Australia
2. School of Ecosystem and Forest Science; University of Melbourne; Parkville 3010 Melbourne Victoria Australia
3. Centre of Excellence for Biosecurity Risk Analysis (CEBRA), School of BioSciences; University of Melbourne; Parkville 3010 Melbourne Victoria Australia
4. Biosis Pty Ltd; Port Melbourne 3207 Melbourne Victoria Australia