1. COWI A/S; Parrallelvej 2 DK-2800 Lyngby Denmark
2. University of Copenhagen, Department of Food and Resource Economics, Center for Macroecology; Evolution and Climate; Rolighedsvej 23 DK-1958 Frederiksberg C Denmark
3. University of Copenhagen, Department of Biology, Center for Macroecology; Evolution and Climate; Universitetsparken 15 DK-2100 Copenhagen O Denmark