1. School of Biological Sciences University of East Anglia Norwich Research Park Norwich NR4 7TJ U.K.
2. Dipartimento di Biologia Università degli Studi di Firenze Via Madonna del Piano 6 50019 Sesto Fiorentino FI Italy
3. Centre for Biodiversity & Environment Research University College London Medawar Building, Gower Street London WC1E 6BT U.K.