1. U.S. Geological Survey, Arkansas Fish and Wildlife Cooperative Research Unit University of Arkansas Fayetteville 72701 AR U.S.A.
2. Department of Integrative Biology Michigan State University East Lansing 48824 MI U.S.A.
3. Department of Biological Sciences University of Arkansas SCEN 522 Fayetteville 72701 AR U.S.A.
4. Wildlife and Research Divisions Arkansas Game & Fish Commission 2 Natural Resources Drive Little Rock 72205 AR U.S.A.