1. South African Institute for Aquatic Biodiversity; Grahamstown 6140 South Africa
2. Department of Ichthyology and Fisheries Science; Rhodes University; Grahamstown 6140 South Africa
3. Institut de Recherche pour le Développement; UMR MARBEC (IRD, Ifremer Univ. Montpellier, CNRS); Avenue Jean Monnet CS 30171 34203 Sète cedex France
4. Aix Marseille Université; Université Toulon, CNRS, IRD, Mediterranean Institute of Oceanography (MIO); Marseille France
5. Port Elizabeth Museum at Bayworld; Port Elizabeth Eastern Cape South Africa
6. Department of Zoology; CMR, Nelson Mandela Metropolitan University; P. O. Box 77000, Port Elizabeth Eastern Cape South Africa
7. Centre d'Etudes Biologiques de Chizé; UMR 7372 CNRS-Université de La Rochelle; 79360 Villiers-en-Bois France