1. Department of Biological and Environmental Science; University of Jyväskylä; Jyväskylä; Finland
2. Department of Biology; Zoological Museum; University of Turku; Turku; Finland
3. Metsähallitus; Natural Heritage Services; Jyväskylä; Finland
4. Metapopulation Research Group; Department of Biosciences; University of Helsinki; Helsinki; Finland