1. Institute of Oceanology; Chinese Academy of Sciences; Qingdao China
2. University of Chinese Academy of Sciences; Beijing China
3. Key Laboratory of Sustainable Development of Marine Fisheries; Ministry of Agriculture; Yellow Sea Fisheries Research Institute; Chinese Academy of Fishery Sciences; Qingdao China
4. Function Laboratory for Marine Fisheries Science and Food Production Processes; Qingdao National Laboratory for Marine Science and Technology; Qingdao China
5. Zhangzidao Sea Ranching Research Center; Dalian China