1. Current Address: Laboratory of Ecology and Evolutionary Biology Yunnan University Kunming 650091 Yunnan China
2. Department of Botany, Prithvi Narayan Campus Tribhuvan University Pokhara Nepal
3. School of Biological Sciences Monash University Melbourne VIC 3800 Australia
4. Faculty of information Technology Monash University Melbourne VIC 3800 Australia
5. School of Media and Communications RMIT University Melbourne VIC 3001 Australia
6. Department of Physiology Monash University Melbourne 3800 Australia