1. Department of Pathology; University of Washington; Seattle WA USA
2. Molecular and Cellular Biology Program; University of Washington; Seattle WA USA
3. Department of Genome Sciences; University of Washington; Seattle WA USA
4. CAS Key Laboratory for Computational Biology; CAS-Max Planck Partner Institute for Computational Biology; Shanghai Institutes for Biological Sciences; Chinese Academy of Sciences; Shanghai China
5. Buck Institute for Age Research; Novato CA USA
6. Institute of Aging Research; Guangdong Medical College; Dongguan China