1. Department of Ecology and Evolutionary Biology; University of Connecticut; Storrs Connecticut CT 06269 U.S.A.
2. Department of Entomology; The Natural History Museum; London SW7 5BD U.K.
3. Swiss Federal Research Institute WSL; Birmensdorf CH-8903 Switzerland
4. Division of Entomology; University of Idaho; Moscow Idaho ID 83844 U.S.A.
5. Department of Ecology and Evolutionary Biology; Yale University; New Haven Connecticut CT 06520 U.S.A.
6. National Institute for Agro-Environmental Sciences; Tsukuba City Ibaraki 305 8604 Japan