1. Key Laboratory of Potato Biology and Biotechnology; Ministry of Agriculture and Rural Affairs; Huazhong Agricultural University; Wuhan 430070 China
2. Key Laboratory of Horticultural Plant Biology (HZAU); Ministry of Education; Huazhong Agricultural University; Wuhan 430070 China
3. College of Horticulture; South China Agricultural University; Guangzhou 510642 China
4. Max Planck Institute of Molecular Plant Physiology; Am Mühlenberg 1 14476 Potsdam-Golm Germany
5. Council for Agricultural Research and Economics; Research Center; CREA-OFA; Via di Fioranello 52 00134 Rome Italy