1. Sorbonne Universités; UPMC Univ Paris 06; UMR 7193; Institut des Sciences de la Terre Paris (iSTeP); F-75005 Paris France
2. CNRS; UMR 7193; Institut des Sciences de la Terre Paris (iSTeP); F-75005 Paris France
3. Natural History Museum of Denmark; Section of Biosystematics; Universitetsparken 15, 2100 Copenhagen Denmark
4. Canterbury Museum; Rolleston Avenue Christchurch 8013 New Zealand
5. Museum of New Zealand Te Papa Tongarewa; PO Box 467 Wellington 6140 New Zealand