1. The James Hutton Institute Aberdeen UK
2. Department of Applied Ecology Fujian Agriculture and Forestry University Fuzhou Fujian China
3. Azorean Biodiversity Group, cE3c ‐ Centre for Ecology, Evolution and Environmental Changes University of the Azores Angra do Heroísmo Portugal
4. School of Biological and Environmental Sciences, James Parsons Building Liverpool John Moores University Liverpool UK
5. Department of Rural Land Use SRUC, Craibstone Estate Aberdeen UK