1. Division of Agricultural and Environmental Sciences; University of Nottingham, Sutton Bonington Campus; Leicestershire; LE12 5RD; UK
2. School of Environmental and Rural Sciences; University of New England; Armidale; New South Wales; 2351; Australia
3. Sustainable Soils and Grassland Systems; Rothamsted Research, West Common; Harpenden; AL5 2JQ; UK