1. Department of Zoology; University of Cambridge; Downing St. Cambridge CB2 3EJ United Kingdom
2. Department of Zoology and Entomology; University of Pretoria; Pretoria 0002 South Africa
3. Science Facilities Department; British Museum of Natural History; Cromwell Road London SW7 5BD United Kingdom
4. Laboratory of Artificial and Natural Evolution; Department of Genetics & Evolution; Sciences III Building; 30 Quai Ernest-Ansermet 1211 Geneva Switzerland
5. Department of Genetics, Evolution, and Environment and Department of Earth Sciences; University College London; Wolfson House to Darwin Building; Gower Street London WC1E 6BT United Kingdom