1. Department of Animal & Plant Sciences; University of Sheffield; Sheffield, S10 2TN United Kingdom
2. Department of Biology; Stanford University; Stanford California 94305 USA
3. Centre for Ecology & Conservation; School of Biosciences; University of Exeter; Penryn Cornwall TR10 9EZ United Kingdom
4. Station d'Ecologie Expérimentale du CNRS USR 2936; 09200 Moulis France