1. National Engineering Research Center for Tree Breeding and Ecological Restoration College of Biological Sciences and Biotechnology Beijing Forestry University Beijing 100083 China
2. Department of Biology School of Life Sciences Institute of Plant and Food Science Southern University of Science and Technology (SUSTech) Shenzhen518055 GuangdongChina
3. The State Key Laboratory for Biology of Plant Disease and Insect Pests Institute of Plant Protection Chinese Academy of Agricultural Sciences Beijing 100193 China
4. International Research Organization for Advanced Science and Technology (IROAST) Kumamoto University Kumamoto 860‐8555 Japan
5. Graduate School of Science and Technology Kumamoto University Kumamoto 860‐8555 Japan