1. School of Biosciences Cardiff University Cardiff CF10 3AX UK
2. Department of Biosciences Durham University Durham DH1 3LE UK
3. The John Bingham Laboratory NIAB Huntingdon Road Cambridge CB3 0LE UK
4. National Research Council of Canada 110 Gymnasium Place Saskatoon SK S79 0W9 Canada
5. Department of Biological and Medical Sciences Oxford Brookes University Oxford OX3 0BP UK