1. Department of Life Sciences, MARE – Marine and Environmental Sciences Centre University of Coimbra Calçada Martim de Freitas 3000‐456 Coimbra Portugal
2. Department of Geography and Tourism, CEGOT – Centre for Studies in Geography and Spatial Planning University of Coimbra Largo da Porta Férrea 3004‐530 Coimbra Portugal
3. Department of Life Sciences, CFE – Centre for Functional Ecology University of Coimbra Calçada Martim de Freitas 3000‐456 Coimbra Portugal