1. Southern Regional Collaborative Innovation Center for Grain and Oil Crops in China, College of Resources and Environmental SciencesHunan Agricultural University Changsha 410128 China
2. Hunan Provincial Key Laboratory of Farmland Pollution Control and Agricultural Resources UseHunan Provincial Key Laboratory of Nutrition in Common University, National Engineering Laboratory on Soil and Fertilizer Resources Efficient Utilization Changsha 410128 China
3. National Key Laboratory of Plant Molecular Genetics and CAS Center for excellence in Molecular Plant SciencesShanghai Institute of Plant Physiology and Ecology, Chinese Academy of Sciences Shanghai 200032 China