1. Department of Grass Science, College of Forestry and Landscape ArchitectureSouth China Agricultural University Guangzhou 510642 China
2. Guangdong Key Laboratory for Innovative Development and Utilization of Forest Plant GermplasmSouth China Agricultural University Guangzhou 510642 China
3. Laboratory of Plant Genetics and DevelopmentNoble Research Institute Ardmore 73401 Oklahoma
4. School of Life SciencesLanzhou University Lanzhou 730000 China