1. Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering School of Life Sciences Fudan University 2005 Songhu Road Shanghai 200438 China
2. Department of Biological Sciences University of Toronto–Scarborough 1265 Military Trail Toronto ON M1C 1A4 Canada
3. State Key Laboratory of Grassland Agro‐ecosystems College of Pastoral Agriculture Science and Technology Lanzhou University 768 Jiayuguan Road West Lanzhou 730020 China
4. Department of Ecology College of Urban and Environmental Sciences Key Laboratory for Earth Surface Processes of the Ministry of Education Peking University 5 Yiheyuan Road Beijing 100871 China
5. Department of Ecology and Evolutionary Biology University of Toronto 25 Wilcocks Street Toronto ON M5S 3B2 Canada