1. SYSU-Alberta Joint Lab for Biodiversity Conservation, State Key Laboratory of Biocontrol, School of Life Sciences, Sun Yat-sen Univ.; Guangzhou 510275 China
2. Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems, South China Botanical Garden, Chinese Academy of Sciences; Guangzhou 510650 China
3. Dept of Renewable Resources; Univ. of Alberta; AB T6G 2H1 Canada
4. State Key Laboratory of Forest and Soil Ecology, Inst. of Applied Ecology, Chinese Academy of Sciences; PO Box 417 Shenyang 110016 China