1. Department of Plant Sciences; University of Oxford; Oxford OX1 3RB UK
2. Australian Research Council Centre of Excellence in Plant Energy Biology; The University of Western Australia; Perth 6009 Australia
3. Faculty of Health and Applied Sciences; University of the West of England; Bristol BS16 1QY United Kingdom
4. Faculty of Agricultural and Nutritional Sciences, Institute of Plant Nutrition and Soil Science; Kiel University; Kiel 24118 Germany
5. School of Biological Sciences; University of Reading; Reading RG6 6AH UK