1. Veterinary Epidemiology, Economics and Public Health GroupRoyal Veterinary CollegeUniversity of LondonHawkshead Lane Hatfield AL9 7TA UK
2. School of Veterinary MedicineCity University of Hong KongTat Chee Avenue Kowloon, Hong Kong SAR
3. Large Animal Research GroupDepartment of ZoologyUniversity of Cambridge Downing Street Cambridge CB2 3EJ UK
4. Mammal Research InstituteUniversity of Pretoria Hatfield, Pretoria South Africa