1. Marine Apex Predator Research Unit (MAPRU) Department of Zoology Institute for Coastal and Marine ResearchNelson Mandela University Port Elizabeth South Africa
2. Institute of Marine Sciences University of California Santa Cruz Santa Cruz CA USA
3. DST‐NRF Centre of Excellence at the FitzPatrick Institute of African Ornithology Nelson Mandela University Port Elizabeth South Africa
4. Centre d’Etudes Biologiques de Chizé UMR 7372 du CNRS‐Université de La Rochelle Villiers‐en‐Bois France