1. Department of Biomathematics; UCLA Medical School; Los Angeles CA 90024 USA
2. Systemic Conservation Biology; Department of Biology; University of Göttingen; Göttingen 37073 Germany
3. Department of Ecology and Evolution; University of Chicago; Chicago IL 60637 USA
4. Department of Ecology and Evolutionary Biology; UCLA; Los Angeles CA 90024 USA
5. Santa Fe Institute; Santa Fe NM 87501 USA