1. Center for Infectious Disease Dynamics & Department of Biology Huck Institute for Life Sciences Pennsylvania State University University Park PA USA
2. Department of Veterinary Population Medicine University of Minnesota St Paul MN USA
3. U.S. Geological Survey Northern Rocky Mountain Science Center Bozeman MT USA
4. Yellowstone Center for Resources Yellowstone National Park WY USA
5. Department of Ecology, Evolution and Behavior University of Minnesota St Paul MN USA