1. CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Mengla China
2. Center of Plant Ecology, Core Botanical Gardens Chinese Academy of Sciences Mengla China
3. College of Biological Resource and Food Engineering Qujing Normal University Qujing China
4. Key Laboratory of Yunnan Province Universities of Qujing Natural History and Early Vertebrate Evolution Qujing China
5. Agriculture and Environment Department Harper Adams University Newport UK
6. Department of Organismic and Evolutionary Biology, Museum of Comparative Zoology Harvard University Cambridge MA USA