1. School of Mathematical Science Yangzhou University Yangzhou China
2. Key Laboratory of Watershed Geographic Sciences Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences Nanjing China
3. School of Life Sciences Nanjing University Nanjing China
4. Department of Ecology, College of Urban and Environmental Science, and Key Laboratory for Earth Surface Processes of the Ministry of Education Peking University Beijing China
5. Department of Agricultural Science University of Helsinki Helsinki Finland