1. Erguna Forest‐Steppe Ecotone Research Station Institute of Applied Ecology Chinese Academy of Sciences Shenyang China
2. University of Chinese Academy of Sciences Beijing China
3. State Key Laboratory of Vegetation and Environmental Change Institute of Botany Chinese Academy of Sciences Beijing China
4. Department of Terrestrial Ecology Netherlands Institute of Ecology (NIOO‐KNAW) Wageningen The Netherlands
5. Section Plant Ecology and Phytochemistry Institute of Biology Leiden University Leiden The Netherlands