1. Department of Biological Sciences Macquarie University Sydney NSW 2109 Australia
2. Department of Physics, Chemistry and Biology (IFM) Linköping University Linköping 58183 Sweden
3. Fowlers Gap Arid Zone Research Station School of Biological, Earth and Environmental Sciences University of New South Wales Sydney NSW 2052 Australia
4. Centre for Ecology and Conservation College of Life and Environmental Sciences University of Exeter, Tremough Campus Penryn TR10 9FE UK