1. Laboratoire de Biométrie et Biologie Evolutive Université Lyon 1, CNRS, UMR 5558 Villeurbanne France
2. Department of Biology University of Louisville Louisville Kentucky USA
3. Department of Biological Sciences and Watershed Studies Institute Murray State University Murray Kentucky USA
4. Institute of Organismic and Molecular Evolution Johannes Gutenberg University Mainz Germany
5. Université Claude Bernard Lyon 1, CNRS, ENTPE, UMR5023 LEHNA Villeurbanne France
6. Rocky Mountain Biological Laboratory Gothic Colorado USA