1. School of Life Sciences; University of Nottingham; Nottingham UK
2. Department of Biology; Aquatic Ecology Unit; Lund University; Lund SE-223 62 Sweden
3. Department of Environmental and Life Sciences; Karlstad University; Karlstad 651 88 Sweden
4. National institute of Aquatic Resources; Technical University of Denmark; Silkeborg DK-8600 Denmark
5. Department of Fish Ecology and Evolution; EAWAG Swiss Federal Institute of Aquatic Science and Technology; Center for Ecology, Evolution and Biogeochemistry; Kastanienbaum Switzerland