1. School of Marine Science and Technology; Newcastle University; Newcastle NE1 7RU UK
2. Institute for Marine Resources and Ecosystem Studies, IMARES; 1790AD Den Burg The Netherlands
3. Centro de Investigación en Ciencias del Mar y Limnología and Escuela de Biología; Universidad de Costa Rica; San Pedro 11501-2060 San José Costa Rica
4. Marine Spatial Ecology Lab; College of Life and Environmental Sciences; University of Exeter; Exeter EX4 4PS UK
5. Marine Spatial Ecology Lab; School of Biological Sciences; University of Queensland; St. Lucia Qld 4072 Australia