1. Department of Biology; West Chester University; 750 S. Church St.; West Chester; PA; USA
2. Department of Zoology; Edward Grey Institute; University of Oxford; South Parks Road; Oxford; OX1 3PS; UK
3. Centre d'Ecologie Fonctionnelle et Evolutive - UMR 5175; Campus CNRS; 1919 Route de Mende; Montpellier Cedex 5; 34293; France