1. Museum of Vertebrate Zoology; University of California; 3101 Valley Life Sciences Building Berkeley California 94720 USA
2. Department of Integrative Biology; University of California; 3060 Valley Life Sciences Building Berkeley California 94720 USA
3. Division of Biological Sciences; University of Montana; Missoula Montana 59812 USA
4. Research School of Biology and Centre for Biodiversity Analysis; Australian National University; Canberra ACT 0200 Australia