1. Department of Molecular and Cellular Physiology; Graduate School of Medicine; Kyoto University; Sakyo-ku Kyoto 606-8501 Japan
2. Department of Obstetrics and Gynecology; Graduate School of Medicine; Osaka University; Suita Osaka 565-0871 Japan
3. Department of Orthopedic Surgery; Graduate School of Medicine; Osaka University; Suita Osaka 565-0871 Japan
4. Cell Biology and Metabolism Group; Graduate School of Frontier Biosciences; Osaka University; Suita Osaka 565-0871 Japan
5. Department of Frontier Biosciences; Graduate School of Frontier Biosciences; Osaka University; Suita Osaka 565-0871 Japan
6. Laboratory of Protein Metabolism; Tokyo Metropolitan Institute of Medical Science; Tokyo 156-8506 Japan
7. Proteo-Science Center; Ehime University; Matsuyama Ehime 790-8577 Japan
8. Department of Biochemistry; Biocenter; University of Würzburg; Am Hubland Würzburg 97074 Germany