1. Department of Ecology and Evolutionary Biology Yale University PO Box 208106 New Haven CT 06520 USA
2. Department of Biology The College of New Jersey 2000 Pennington Rd Ewing NJ 08628 USA
3. Division of Botany Peabody Museum of Natural History Yale University PO Box 208118 New Haven CT 06520 USA
4. Laboratorio de Botánica y Sistemática Departamento de Ciencias Biológicas Universidad de los Andes Apartado Aéreo 4976 Bogotá Colombia
5. Department of Ecology and Evolutionary Biology Brown University Box G‐W 80 Waterman Street Providence RI 02912 USA