1. Applied Behavioural Ecology and Ecosystem Research Unit (ABEERU), Department of Environmental Sciences, College of Agriculture and Environmental Sciences University of South Africa Florida Gauteng South Africa
2. Molecular Ecology and Evolution Programme, Department of Biochemistry, Genetics and Microbiology, Faculty of Natural and Agricultural Science University of Pretoria Pretoria Gauteng South Africa
3. South African National Biodiversity Institute Pretoria South Africa
4. School of Health and Life Sciences Teesside University Middlesbrough UK